From that article.

Dire wolves are iconic beasts. Thousands of these extinct Pleistocene carnivores have been recovered from the La Brea Tar Pits in Los Angeles. And the massive canids have even received some time in the spotlight thanks to the television series Game of Thrones. But a new study of dire wolf genetics has startled paleontologists: it found that these animals were not wolves at all, but rather the last of a dog lineage that evolved in North America.

This makes me wonder if the Carolina Dog and probably the Dingo bear any DNA similarity to the Dire Wolf